Formal updated conservation advice is now available for Darwin Mounds SAC under the conservation advice tab below.


Darwin Mounds MPA


Status: Special Area of Conservation (SAC)Click to link to the interactive map


Darwin Mounds SAC lies at the north end of the Rockall Trough located approximately 160 km north west of Cape Wrath, Scotland, with depths between 710-1129m.


The site contains an extensive area of sandy mounds, each capped with multiple thickets of cold-water corals. Unlike most cold-water corals which attach to hard-surfaces, at this site the corals is growing on sand. The thickets of coral range in size from one to several metres in diameter and support many other species, such as starfish, sponges and large populations of xenophyophores. Xenophyophores are the world’s largest single celled organisms and are only found in deep-sea environments. They provide shelter for a range of small sea creatures, many of which are prey for larger species, and act as an elevated perch for filter feeders such as brittlestars.


More detailed site information can be found on the Summary tab.


Map displaying MPA boundary and

associated protected feature data.

Visit the JNCC MPA Mapper to further

view and explore data for this MPA.


Legislation behind the designation: EU Habitats Directive 1992 transposed into UK law by the Conservation of Offshore Marine Habitats and Species Regulations 2017.


Protected features

Features Feature Type
1170 Reefs Annex I Habitat*

* For the latest Annex I habitat resource figures, please see the link to the latest Habitats Directive Article 17 reporting in the Assessment tab.


The acquisition of new data may result in updates to our knowledge on feature presence and extent within this site. The most up to date information is reflected on the map at the top of the page and in JNCCs MPA mapper, with the evidence underpinning available in the Evidence tab.

Site Timeline

The diagram below summarises the key milestones involved in the selection and designation of Darwin Mounds SAC.  More detail can be found within the Relevant Documentation.

Darwin Mounds Timeline

Relevant Documentation

The documents referred to below and any other historical documents relating to Darwin Mounds SAC were produced during the selection and designation process and therefore may be out of date.  This site information centre is the most up to date source of information for Darwin Mounds SAC, and will reflect any additional information gathered since these documents were produced. Information about the SAC site selection process is also available on the JNCC SAC pages.




Last updated: October 2017


Information for this site summary was adapted from the SAC selection assessment and incorporates any further information gathered since this document was produced. Please refer to this document for further details and information sources.


Site overview: The Darwin Mounds is located approximately 160km north-west of Cape Wrath, Scotland at the north end of the Rockall Trough. The site consists of a large number of coral topped ‘sand volcanoes’ composed of sand overlying mud. The individual mounds are up to 75m wide and 5m high and are morphologically unique in UK waters. The mounds themselves are unusual geological features that are thought to have been formed by fluid expulsion from the seabed. The mounds are most elevated to the north of the site and diminish in height to the south. There are two denser regions of mounds in the north-east and north-west of the site. The north-east field of the Darwin Mounds measures approximately 13km by 9km and contains approximately 150 mounds, whereas the north-west field measures approximately 13km by 4km and contains approximately 75 mounds.


The mounds in the northern part of the site showed an acoustic signature of a “tail” which generally lies to the south west of the mound feature consisting of elongate to oval patches up to 500m long. The 'tail' feature of the mounds is thought to be unique globally. The tails are not topographically distinct from the surrounding seabed but are covered with a fine veneer of sand. They support substantial numbers of the xenophyophore, Syringammina fragilissima (a single celled organism that can grow up to 20cm in diameter), in densities greater than the surrounding seabed.


On the majority of the mounds there are also multiple colonies of Lophelia pertusa growing, which in many cases have formed thickets which arise from the surface of the mound. This site was the first observed record of cold-water coral reefs growing on sand at the time of discovery. Like the topography of the mounds, the frequency of coral communities diminishes towards the south of the mound field. Although the reef habitat on top of the mounds is formed primarily from Lophelia pertusa another cold-water coral, Madrepora oculata, is also present. The thickets of cold-water corals provide a habitat for echiuran worms, brittlestars, brisingiid starfish and sponges. Various fish have been observed among the mounds, but not at higher densities than the wider environment.


Within the Atlantic north-west approaches, Rockall Trough and Faroe-Shetland Channel Regional Sea where Darwin Mounds SAC is situated, there are three other SACs designated for the presence of Annex I reef: Anton Dohrn SAC, North West Rockall Bank SAC and East Rockall Bank SAC. Each site represents different types of Annex I reef. Wyville Thomson Ridge SAC on the edge of the adjacent Scottish continental shelf regional sea is also designated for Annex I reef. The tail feature observed on mounds in the northern part of the Darwin Mounds site is thought to be globally unique from a geological perspective. Further detail on the evidence for this SAC can be found on the Evidence tab.


Site location:  Coordinates for this SAC can be found in the standard data form listed in the relevant documentation.

Site area:  1,377 km2, a similar size to the Brecon Beacons National Park (1,344 km2)

Site depth range:  710 – 1,129m.

Charting Progress 2 Biogeographic Region: Atlantic North-West approaches, Rockall Trough and Faroe-Shetland Channel. A small part of the site extends into the Scottish continental shelf biogeographic region.


Site boundary description

The boundary of the Darwin Mounds SAC is a simple polygon, fully enclosing the minimum area necessary to ensure protection of the Annex I feature. The boundary has been aligned with the area permanently closed to bottom trawling through the European Commission common fisheries policy (EC Regulation 602/2004).

As bottom trawling is a significant threat to the interest feature the proposed boundary includes a margin to ensure its protection. The maximum depth of water around the feature is 1,000-1,100m. Fishing vessels which are bottom trawling in the region need a minimum towline length of twice the depth of water in which they are fishing. Assuming a ratio of 2:1 fishing warp length to depth, the proposed boundary is defined to include a margin of 2,200m from the cold-water coral reefs.




Last updated: October 2017


For a full overview of the data used to support site identification along with information on confidence in feature presence and extent see the Darwin Mounds SAC selection assessment. Data for this SAC have been primarily collected through JNCC-funded or collaborative surveys with other data obtained through other data sourcing.  The data gathered provide direct evidence confirming the presence of the protected features within the site. Additional survey data will be added to JNCCs MPA mapper in due course.


Survey and data gathering

  • Deep Links Project (2016) - A collaborative project between Plymouth University's Deep Sea CRU and University of Oxford, in partnership with JNCC and British Geological Survey, funded by NERC. During May and June 2016 the team undertook a 6 week research cruise in the North East Atlantic, including Anton Dohrn Seamount, on board the RRS James Cook collecting data. This project aims to investigate the theory that populations at bathyal depths are more isolated because the currents that transport larvae decrease with depth.
  • Survey of Darwin Mounds and North West Rockall (2011)
    JNCC collaborated with the National Oceanography Centre and Plymouth University to carry out habitat mapping to assess the status of different benthic habitats in relation to human activities, especially deep-sea bottom trawling. Acoustic and underwater imagery data were collected.
  • RRS Charles Darwin Cruise 123C3-4 (2000)
    The cruise formed part of the continuing AMES undertaken by the National Oceanography Centre in waters to the north and west of Scotland. Seabed samples and underwater imagery were collected to study environmental parameters and benthic communities.
  • RRS Charles Darwin cruise 119C leg B (1999)
    The cruise was undertaken by the National Oceanography Centre and expanded on the work undertaken on the AFEN Atlantic Margin Environmental Survey (AMES). The cruise collected acoustic maps of the seafloor, along with photographic and video observations. Where possible, seabed samples were also collected to analyse selected environmental parameters.
  • Atlantic Frontier Environmental Network Survey (1998)
    The Atlantic Frontier Environment Network (AFEN; including JNCC) commissioned a survey to map and sample the seabed to the north and west of Scotland, allocated for oil and gas exploration in 1997. The aim of the survey was to provide the shape and texture of the seabed, the fauna and the physical processes affecting them, both of today and over the last 10-15,000 years. The Darwin Mounds were discovered on this cruise.


Data analysis reports
Further analysis of data gathered as part of the surveys listed above are available via the following reports:

Additional relevant literature
References for further supporting scientific literature consulted during the identification of this site can be found in the site selection assessment. Please be aware that although these sources contain information which is of interest in relation to this MPA, they do not necessarily represent the views of JNCC:

  • Huvenne V.A.I., Bett B.J., Masson D.G., Le Bas T.P. and Wheeler A.J. (2016) Effectiveness of a deep-sea cold-water coral Marine Protected Area, following eight years of fisheries closure. Biological Conservation, 200: 60–69.
  • Masson, D.G., Bett, B.J., Billett, D.S.M., Jacobs, C.L., Wheeler, A.J. and Wynn, R.B., (2003) The origin of deep-water, coral-topped mounds in the northern Rockall Trough, Northeast Atlantic. Marine Geology, 194: 159-180.
  • Bett, B.J. (2001) UK Atlantic Margin Environmental Survey: Introduction and overview of bathyal benthic ecology. Continental Shelf Research, 21: 917-956.
  • Bett, B.J., Billett, D.S.M., Masson, D.G., Tyler, P.A. et al. (2001) RRS Discovery Cruise 248: 07 Jul-10 Aug 2000. A multidisciplinary study of the environment and ecology of deep-water coral ecosystems and associated seabed facies and features (The Darwin Mounds, Porcupine Bank and Porcupine Seabight). National Oceanography Centre Cruise Report 36: 108. This cruise formed part of the work done by the National Oceanography Centre for the EU funded Atlantic Coral Ecosystem Study and ECOMOUND projects. These projects  investigated the geological processes underlying the formation of the Darwin Mounds, and studied the biology and ecology of cold water coral reefs.


Knowledge gaps
If you are aware of any additional data or relevant scientific papers for this site not listed here, please contact JNCC.


Conservation Advice


Last updated: March 2018

This is the formal conservation advice for Darwin Mounds SAC.  Further information on the approach used to develop this advice is available on the Conservation Advice webpage

You must refer to this advice if you:

  • Undertake a Habitats Regulation Assessment (HRA) for a plan or project that could impact the site;
  • Provide information for a HRA;
  • Respond to specific measures to prevent the deterioration of habitats and significant disturbances of features; and
  • Consider the need to put new or additional management measures in place.


You may also find it useful to refer to this advice if you:

  • Carry out any other activity that could impact the site.


We will engage with stakeholders to identify any lessons which JNCC can learn from customers who have used the advice, with a view to continuing to ensure it is fit-for-purpose.


The following table provides an overview of the components of the conservation advice, and provides hyperlinks to each of the products for this MPA. These elements together form JNCC’s formal conservation advice for this site and should be read in conjunction with each other. This advice reflects the most up-to-date evidence held by JNCC (correct as of March 2018). A zipped folder enabling these documents to be downloaded together is available at the bottom of this page. 


Document Overview
Background Information Explains the purpose of the advice and when it must be referred to.
Conservation Objectives


Supplementary Advice on the Conservation Objectives (SACO)

The Conservation Objectives set out the broad ecological aims for the site. JNCC provide supplementary advice in the SACO which is essential reading to support interpretation of these conservation objectives.


You can use these documents to assess the impacts of your planned activity on the important attributes of the site.


Please note our current understanding of whether the available evidence indicates that each attribute needs to be restored or maintained is not provided here. However, links to available evidence for the site are provided and should you require further site-specific information for the site, please contact JNCC at:


Conservation Advice Statements

These statements provide a summary of the Supplementary Advice on the Conservation Objectives (SACO).


  • Site condition presents our up to date understanding of the condition of features within the site;
  • Conservation benefits which the site can provide, these help you understand what is important about the site and why it needs protecting; and
  • Conservation measures which JNCC consider are needed to support achievement of the conservation objectives. These provide clarity around measures needed to support restoration or maintenance of the feature(s) within the site.
Advice on Operations

Provides information on the activities capable of affecting site integrity and therefore achievement of the site’s conservation objectives.


This is a starting point for determining potential management requirements. It does not take into account the intensity, frequency or cumulative impacts from activities taking place. It is simply to advise you of the possible adverse impacts that your activity can have on a MPA’s features.


Use the advice on operations to determine those pressures your activity causes that could harm the habitat and/or species features of the site.




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Activities and Management


Last updated: October 2017


Management status: Progressing towards being well managed


Information about the long-term recovery of cold-water coral populations is scarce but fisheries management measures are in place. Continued protection of the Darwin Mounds area combined with regular monitoring is necessary to obtain further insights in these processes (Howell et al 2014).

  1. The documentation of appropriate management information - conservation objectives, advice on activities capable of affecting the protected features of a site, and spatial information on the presence and extent of the protected features of a site.
  2. The implementation of management measures - management actions considered necessary to achieve the conservation objectives of a site.
  3. Site condition monitoring programmes – collecting the information necessary to determine progress towards a sites conservation objectives.
  4. Assessment of progress towards conservation objectives – using available information to infer whether or not a site is moving towards or has achieved its conservation objectives.
  • The conservation objectives and advice on activities capable of affecting the conservation status of the protected feature of this site are available under the conservation advice tab.
  • JNCC are in the process of improving our MPA conservation advice packages. Further information is available on our conservation advice pages.
  • Spatial information on the presence and extent of the protected feature of this MPA is available via JNCCs MPA mapper.
  • JNCC are in the process of developing downloadable MPA data packages where appropriate permissions to share datasets are in place.


Licensable activities

  • Whilst ‘licensable’ activities such as oil and gas exploration and production do not take place within Darwin Mounds SAC at present, any future proposals would have to comply with Article 6 (3) of the EU Habitats Directive 1992, which is transposed into UK law by the Conservation of Offshore Marine Habitats and Species Regulations 2017.
  • Our conservation advice supports the consents process by setting out the conservation objectives for the protected feature of this MPA and advice on activities that may result in pressures to which the protected feature is considered sensitive.
  • Further information on JNCCs role in the provision of advice for licensed activities in the UK offshore area is available on JNCCs offshore industries advice webpage


Site condition monitoring

Fishing vessel monitoring data is used to monitor compliance with the management measure in place. A baseline condition monitoring survey was undertaken in 2011 (reported in Howell et al 2014). Further information is provided in the Monitoring tab.


Assessment of progress towards conservation objectives

No long-term condition monitoring data is available to determine whether the MPA is moving towards or has reached its conservation objectives. Howell et al (2014) suggest that ecosystem recovery is still a long way off, as no signs of coral recolonisation and limited coral regrowth were recorded in damaged areas after eight years of closure to bottom-contacting fishing gear (since 2004). Further information is provided in the Assessment tab.




Last updated: October 2017


JNCC is currently leading on the development of a strategy for biodiversity monitoring across all UK waters, to include SAC monitoring. For SACs, data and evidence collected from monitoring activities will aim to:

  • Enable assessment of condition of the features within sites;
  • Enable assessment of the degree to which management measures are effective in achieving the conservation objectives for the protected features;
  • Support the identification of priorities for future protection and/or management; and
  • Enable the government to fulfil its national and international assessment and reporting commitments in relation to MPAs and help identify where further action may be required.


Information on monitoring of this SAC will be provided when it becomes available.




Last updated: October 2017


Assessments of the condition of designated features in offshore MPAs are required to report against our legal obligations. Ideally these assessments are based on observed data, and then measured against targets for predefined indicators. However, for MPAs in offshore waters the appropriate information is not always available particularly for seabed habitats, which are the main type of designated features within offshore MPAs. 


To address these challenges, JNCC has been an active partner in developing new approaches and tools for the assessment of habitats and species for a variety of national and international status reports. These tools cover methods for producing interim assessments of site features and their responses to pressures, as well as developing more robust indicators for determining condition of the features. The reports include the second cycle of the conservation status assessment reports under the EU Habitats Directive, Charting Progress 2 and the OSPAR Quality Status Report. JNCC continues to develop and pilot tools for the assessment of marine habitats and species in offshore waters to improve the quality and transparency of our assessments, and contribute to the monitoring of marine biodiversity in UK waters.


Every six years, member states are also required under Article 17 of the EU Habitats Directive to report on the conservation status of Annex I habitats and Annex II species on the Habitats Directive.  The assessments should consider the habitat or species both within the Natura 2000 network and in the wider sea.  The latest report was submitted by the UK in 2013 and provided a second assessment of the conservation status of relevant habitats and species within UK marine waters during 2007-2012. The next report is for the period 2013-2018 and is due in 2019; information on the condition of features within SACs will make a contribution to this report.  These assessments will also feed into six yearly reports on the state of the marine environment under the Marine Strategy Framework Directive, which aims to achieve good environmental status by 2020.



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