Lowland neutral meadows and pastures consist of a rich mixture
of native grasses and broad-leaved herbs. They occur throughout
lowland UK, often on shallow slopes or level ground with relatively
deep soils that is neither strongly acidic nor lime-rich. The
meadows may be managed for hay cropping, usually with grazing of
the aftermath (vegetation that re-grows following cutting), or by
grazing as permanent pasture.
Up to 35 or more plant species may occur in a 2x2m sample,
including grasses such as crested dog’s tail Cynosurus cristatus
and red fescue Festuca rubra, and herbs such as knapweed
Centaurea nigra, bird’s-foot trefoil Lotus corniculatus and ox-eye
daisy Leucanthemum vulgare. Some pastures may be important
for waxcap and earth-tongue fungi. Old meadows and pastures can
support a rich insect community, including butterflies,
grasshoppers, bumblebees and yellow meadow ants. They can also
provide important feeding areas for birds such as the linnet
Carduelis cannabina and meadow pipit Anthus pratensis, and
bats and small mammals such as the field vole Microtus
agrestis.
The flora of lowland meadows can include rare and scarce species
such as snakes’s head fritillary Fritillariameleagris,
Sulphur clover Trifolium
ochroleucon, field gentian Gentianella campestris and
green-winged orchid Orchis morio. This may be matched
by a scarce invertebrate fauna, including hornet
robber-fly Asilus crabroniformis and
shrill carder bee Bombus silvarum.
Lowland meadows also include the now scarce flood-meadows of
central England and eastern Wales, which rely on seasonal flooding
in winter, and support tall, moisture-loving species such as great
burnet Sanguisorba officinalis, meadowsweet Filipendula
ulmaria and pepper-saxifrage Silaum silaus.
Upland hay meadows
Upland or mountain hay meadows are a distinctive, rare habitat
of northern England and central Scotland. They occur in scattered
fields or in small isolated groups of fields in the upland fringes.
The hay meadows are typically grazed by cattle or sheep from the
autumn through to early spring, before being ‘shut-up’ to allow the
hay crop to grow. Hay is normally cut in July or August, but in
some cases as late as September. The livestock are then typically
returned to the field to graze the aftermath (vegetation that
re-grows following cutting). Traditionally, the meadows were
given a light dressing of farmyard manure in the spring, together
with occasional liming to maintain neutral soil pH conditions.
Upland hay meadows consist of a rich mixture of native grasses
and broad-leaved herbs, including for example wood
crane's-bill Geranium sylvaticum, great burnet
Sanguisorba officinalis and lady's mantle Alchemilla
spp. They are also important as feeding areas for
invertebrates and bats, and provide nesting and feeding sites for
birds such as yellow wagtail, twite, and curlew. Threatened species
of upland hay meadows include various species of rare lady’s mantle
Alchemilla, northern hawk’s-beard Crepis mollis
and small-white orchid Leucorchis albida on hay meadow
banks.
Lowland calcareous grassland
Lowland calcareous grasslands are characterised by lime-loving
plants and are found mainly, but not entirely, in the south and
east of the UK, where they occur on shallow, calcareous soils
generally overlying limestone rocks, including chalk. These
grasslands are now largely found on distinct topographic features
such as escarpments or dry valley slopes but occasionally remnants
on flatter topography survive such as on Salisbury Plain or in
Breckland. By contrast, the blue moor-grass Sesleria
grasslands are found up to the upland fringes in northern England.
Calcareous grasslands are typically managed by grazing but may
sometimes be cut for hay.
The flora can be very rich including many nationally rare and
scarce species such as monkey orchid Orchis simia, hoary
rockrose Helianthemum canum and pasque flower
Pulsatilla vulgaris. This can be matched by an equally
diverse invertebrate fauna including scarce species like the Adonis
blue Lysandra bellargus and the wart-biter cricket
Decticus verrucivorus. These grasslands also provide
feeding and breeding habitat for birds such as the stone curlew
Burhinus oedicnemus and skylark Alauda
arvensis.
Lowland dry acid grassland
Lowland
acid grasslands are widely distributed throughout the UK, typically
on nutrient-poor, acid soils overlying sands and gravels, hard
volcanic rocks or sandstones. Large areas occur in the upland
fringes (generally below 300m) but more important are the
well-drained often parched habitats that are found in the drier
lowland areas, such as the East Anglian Breckland, the New Forest
and the Weald. Species-rich forms are also very occasionally found
in the western hills. Lowland acid grassland is often an important
component of heathland landscapes, of old parklands and of commons,
and more locally, of coastal cliffs and shingle. It is normally
managed as pasture.
Species richness can be variable in lowland acid grassland,
ranging from less than 5 to greater than 25 plant species per 2x2m
sample. Parched acid grassland can provide habitat for a number of
rare and scarce plants including mossy stonewort Crassula
tillaea, sticky catchfly Lychnis viscaria and spring
speedwell Veronica verna. The open, sandy soils can
support considerable numbers of ground-dwelling and burrowing
invertebrates such as solitary bees and wasps, and are favoured by
birds such as the stone-curlew Burhinus oedicnemus,
woodlark Lullula arborea and nightjar Caprimulgus
europaeus. By contrast, fungi such as wax-caps, earth-tongues
and fairy clubs favour grazed, moist acid grassland, particularly
in the west of the UK.
Purple moor grass and rush pasture
The marshy grasslands known as purple moor grass and rush
pastures traditionally provided useful grazing in dry summers. They
are found throughout the UK, although are mainly concentrated in
high rainfall areas of the west on gently sloping land with
poorly-drained soils. Important areas are the Culm grasslands of
Devon and Cornwall, the Rhôs pastures of Wales and the fen meadows
of County Fermanagh. Today they are still used as rough grazing for
cattle or ponies with an occasional hay crop, although increasingly
management is being abandoned altogether.
They are usually dominated by purple moor-grass Molinia caerulea
and/or rushes, especially sharp-flowered rush Juncus
acutiflorus, but may include up to 50 plant species in a 2x2m
sample including characteristic species such as meadow thistle
Cirsium dissectum and whorled caraway Carum verticillatum. The
habitat supports a varied invertebrate fauna; the best known
example being the marsh fritillary butterfly Euphydryas
aurinia. Curlew Numenius arquata and lapwing Vanellus
vanellus breed in marshy pastures and other species using the
habitat include snipe Gallinago gallinago, barn owl Tyto alba,
grass snake Natrix natrix and adder Vipera berus.
Calaminarian grassland
Calaminarian grasslands can be found on soils
that have levels of heavy metals, such as lead, zinc chromium and
copper, which are toxic to most plant species. They mostly occur in
the north and west of the UK on artificial habitats resulting from
past mining activity, although there are a few near-natural
examples. The habitat occurs in three main situations: as
near-natural, open vegetation of serpentine rock and mineral vein
outcrops with skeletal soils, such as at the Keen of Hamar in
Shetland; on stable river gravels rich in lead and zinc, such as on
the Tyne and Allen river gravels in Northumberland; and on
artificial mine workings and spoil heaps, such as on the Halkyn
Mountain in north Wales.
The vegetation is usually species-poor, but
often includes spring sandwort Minuartia verna and alpine
penny-cress Thlaspi caerulescens. There is a genetically
adapted range of other species including sheep’s fescue Festuca
ovina, sea campion Silene uniflora and thrift
Armeria maritima. The heavy metal toxicity of the soils
combined with paucity of nutrients results in open, slow-growing
vegetation. Rarer species such as forked spleenwort Asplenium
septentrionale can benefit from lack of competition from
vigorous colonists. Some sites hold important populations of rare
bryophytes and lichens.